verified source for familyBell, N. E.; Quandt, D.; O'Brien, T. J.; Newton, A. E. (2007). Taxonomy and phylogeny in the earliest diverging pleurocarps: square holes and bifurcating pegs. <em>The Bryologist.</em> 110(3): 533-560., available online athttps://doi.org/10.1639/0007-2745(2007)110[533:tapite]2.0.co;2[details]