Ratcliffe, B. C. (2014). A new genus and species of Dynastinae (Coleoptera: Scarabaeidae) from the Galápagos Islands, Ecuador, other new species of Cyclocephalini, Pentodontini, and Phileurini from South America, and a revised key to the genera of New World Pentodontini. The Coleopterists Bulletin. 68(4): 663-680.
A new genus and species of Dynastinae (Coleoptera: Scarabaeidae) from the Galápagos Islands, Ecuador, other new species of Cyclocephalini, Pentodontini, and Phileurini from South America, and a revised key to the genera of New World Pentodontini