Challands, T. J.; Smithson, T. R.; Clack, J. A.; Bennett, C. E.; Marshall, J. E. A.; Wallace-Johnson, S. M.; Hill, H. (2019). A lungfish survivor of the end-Devonian extinction and an Early Carboniferous dipnoan radiation. Journal of Systematic Palaeontology. 17(21): 1825-1846.