Nesbitt, S. J.; Denton, R. K.; Loewen, M. A.; Brusatte, S. L.; Smith, N. D.; Turner, A. H.; Kirkland, J. I.; McDonald, A. T.; Wolfe, D. G. (2019). A mid-Cretaceous tyrannosauroid and the origin of North American end-Cretaceous dinosaur assemblages. Nature Ecology & Evolution. 3(6): 892-899.