Ratcliffe, B.C.; Smith, D.M.; Erwin, D. (2005). Oryctoantiquus borealis, new genus and species from the Eocene of Oregon, U.S.A., the world's oldest fossil dynastine and largest fossil scarabaeid (Coleoptera: Scarabaeidae: Dynastinae). Coleopterists Bulletin, 59 (1), 127-135
4497
Ratcliffe, B.C.; Smith, D.M.; Erwin, D.
2005
Oryctoantiquus borealis, new genus and species from the Eocene of Oregon, U.S.A., the world's oldest fossil dynastine and largest fossil scarabaeid (Coleoptera: Scarabaeidae: Dynastinae).