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IRMNG taxon details

Tingidae Laporte, 1833

103421  (urn:lsid:irmng.org:taxname:103421)

  1. Biota
  2. Animalia (Kingdom)
  3. Arthropoda (Phylum)
  4. Hexapoda (Subphylum)
  5. Insecta (Class)
  6. Hemiptera (Order)
  7. Tingidae (Family)
accepted
Family
Tingididae · unaccepted

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Children Display

  1. Genus Acalypta Westwood, 1840
  2. Genus Acanthocheila Stål, 1858
  3. Genus Acanthomoplax Souma & Kamitani, 2021
  4. Genus Acysta Champion, 1898
  5. Genus Agramma Stephens, 1829
  6. Genus Allocader Drake, 1950
  7. Genus Alloeocysta Drake, 1961
  8. Genus Alveotingis Osborn & Drake, 1916
  9. Genus Ambarcader Perrichot, Nel, Guilbert & Néraudeau, 2006 †
  10. Genus Amberobyrsa Heiss, 2009 †
  11. Genus Archetingis Montagna, Strada, Dioli & Tintori, 2018 †
  12. Genus Atheas Champion, 1898
  13. Genus Australotingis Hacker, 1927
  14. Genus Birabena Drake & Hurd, 1945
  15. Genus Bunotingis Drake, 1948
  16. Genus Burmacader Heiss & Guilbert, 2013 †
  17. Genus Burmavianaida Souma, Yamamoto & Takahashi, 2021 †
  18. Genus Caledoderus Guilbert, 2012
  19. Genus Caledoderus Guilbert, 2011
  20. Genus Callithrincus Horváth, 1925
  21. Genus Caloloma Drake & Bruner, 1924
  22. Genus Calotingis Drake, 1918
  23. Genus Cantacader Amyot & Serville, 1843
  24. Genus Carinacader Knudson in Knudson et al., 2020
  25. Genus Carldrakeana Froeschner, 1968
  26. Genus Celantia Distant, 1903
  27. Genus Ceratocader Drake, 1950
  28. Genus Ceratotingis Montemayor, 2008
  29. Genus Cercotingis Shofner & Cassis, 2024
  30. Genus Cervantestingis Henry & Brailovsky, 2018
  31. Genus Chorotingis Drake, 1961
  32. Genus Codotingis Drake, 1942
  33. Genus Compseuta Stål, 1873
  34. Genus Corythaica Stål, 1873
  35. Genus Corythucha Stål, 1873
  36. Genus Cromerus Distant, 1902
  37. Genus Cucullitingis Du & Yao, 2018
  38. Genus Cysteochila Stål, 1873
  39. Genus Dichocysta Champion, 1898
  40. Genus Dictyla Stål, 1874
  41. Genus Dictyonota Curtis, 1827
  42. Genus Dicysta Champion, 1897
  43. Genus Diplocysta Horváth, 1925
  44. Genus Engynoma Drake, 1942
  45. Genus Epimixia Kirkaldy, 1908
  46. Genus Eritingis Drake & Ruhoff, 1962
  47. Genus Esocampylia Hacker, 1929
  48. Genus Euaulana Drake, 1946
  49. Genus Froggattia Froggatt, 1901
  50. Genus Furcilliger Horváth, 1925
  51. Genus Galeatus Curtis, 1833
  52. Genus Gargaphia Stål, 1862
  53. Genus Gyaclavator Wappler, Guilbert, Wedmann, Labandeira in Wappler, Guilbert, Labandeira, Hörnschemeyer & Wedmann, 2015 †
  54. Genus Gymnotingis Hacker, 1928
  55. Genus Henryvianaida Guidoti, Itzel Montemayor, Campos & Guilbert, 2019
  56. Genus Hesperotingis Parshley, 1917
  57. Genus Hypsipyrgias Kirkaldy, 1908
  58. Genus Inoma Hacker, 1927
  59. Genus Inonemia Drake, 1942
  60. Genus Ischnotingis Horvath, 1925
  61. Genus Ittolemma Symonds & Cassis, 2014
  62. Genus Kzylcader Popov & Golub, 2019 †
  63. Genus Lasiacantha Stål, 1873
  64. Genus Latidorsum Wang, Tang & Yao in Wang et al., 2021 †
  65. Genus Leptobyrsa Stål, 1873
  66. Genus Leptodictya Stål, 1873
  67. Genus Leptopharsa Stål, 1873
  68. Genus Leptoypha Stål, 1873
  69. Genus Lepturga Stål, 1873
  70. Genus Limnostatua Souma & Ishikawa, 2022
  71. Genus Lutetiacader Wappler, 2006 †
  72. Genus Majusculispinatus Mu, Ren & Yao in Mu et al., 2025 †
  73. Genus Malagasotingis Lis, 2009
  74. Genus Malandiola Horváth, 1926
  75. Genus Mecopharsa Drake, 1953
  76. Genus Melanorhopala Stål, 1873
  77. Genus Mexicotingis Henry, Montemayor & Knudson, 2017
  78. Genus Nethersia Horváth, 1925
  79. Genus Niborskiana Montemayor, 2012
  80. Genus Oncophysa Stål, 1873
  81. Genus Pampacader Carpintero & Montemayor, 2005
  82. Genus Paraceratotingis Henry, Montemayor & Knudson, 2017
  83. Genus Paracopium Distant, 1902
  84. Genus Parada Horvàth, 1925
  85. Genus Paraphatnomacader Guilbert & Heiss, 2019 †
  86. Genus Parasinalda Heiss & Golub, 2013
  87. Genus Perissonemia Drake & Poor, 1937
  88. Genus Phatnoma Fieber, 1844
  89. Genus Phymacysta Monte, 1942
  90. Genus Physatocheila Fieber, 1844
  91. Genus Pontanus Distant, 1902
  92. Genus Proteatingis Shofner & Cassis, 2024
  93. Genus Pseudacysta Blatchley, 1926
  94. Genus Radinacantha Hacker, 1929
  95. Genus Raunotingis Duarte Rodrigues, 1980
  96. Genus Spinitingis Heiss & Guilbert, 2013 †
  97. Genus Stephanitis Stål, 1873
  98. Genus Swaustraltingis Moir & Guilbert, 2012
  99. Genus Tanybyrsa Drake, 1942
  100. Genus Taurcader Knudson in Knudson et al., 2020
  101. Genus Teleonemia Costa, 1864
  102. Genus Tigavaria Drake, 1946
  103. Genus Tingiphatnoma Guilbert & Heiss, 2019 †
  104. Genus Tingis Fabricius, 1803
  105. Genus Ulonemia Drake & Poor, 1937
  106. Genus Urentius Distant, 1903
  107. Genus Viktorgolubia Popov, 2019 †
  108. Genus Acanthochila Stål, 1873 accepted as Acanthocheila Stål, 1858
  109. Genus Amaurosterphus Stål, 1868 accepted as Teleonemia Costa, 1864
  110. Genus Americia Stål, 1873 accepted as Teleonemia Costa, 1864
  111. Genus Ayrerus Distant, 1903 accepted as Urentius Distant, 1903
  112. Genus Birgitta Lindberg, 1927 accepted as Leptoypha Stål, 1873
  113. Genus Bredenbachius Distant, 1903 accepted as Cysteochila Stål, 1873
  114. Genus Cadamustus Distant, 1903 accepted as Stephanitis Stål, 1873
  115. Genus Calliphanes Horváth, 1906 accepted as Stephanitis Stål, 1873
  116. Genus Campylostira Fieber, 1860 (Misspelling)
  117. Genus Catoplatys Agassiz, 1846 (Unjustified (?) emendation)
  118. Genus Copius Burmeister, 1835 (Misspelling)
  119. Genus Corythuca Lethierry & Severin, 1896 accepted as Corythucha Stål, 1873
  120. Genus Derophysa Douglas & Scott, 1876 (Misspelling)
  121. Genus Derophysia Spinola, 1837 (Misspelling)
  122. Genus Drakea Schouteden, 1953 accepted as Agramma Stephens, 1829
  123. Genus Dyctinota Costa, 1847 accepted as Dictyonota Curtis, 1827
  124. Genus Esocamphylia Drake, 1942 accepted as Esocampylia Hacker, 1929
  125. Genus Gelchossa Kirkaldy, 1904 accepted as Leptopharsa Stål, 1873
  126. Genus Hanuala Kirkaldy, 1905 accepted as Leptodictya Stål, 1873
  127. Genus Jannaeus Distant, 1909 accepted as Lasiacantha Stål, 1873
  128. Genus Leptostyla Stål, 1873 accepted as Leptopharsa Stål, 1873
  129. Genus Lisia Koçak & Kemal, 2010 accepted as Agramma Stephens, 1829
  130. Genus Maecenas Kirkaldy, 1904 accepted as Stephanitis Stål, 1873
  131. Genus Microtingis Heiss & Golub, 2021 † accepted as Burmavianaida Souma, Yamamoto & Takahashi, 2021 †
  132. Genus Mokanna Distant, 1910 accepted as Stephanitis Stål, 1873
  133. Genus Myrmecotingis Hacker, 1928 accepted as Lasiacantha Stål, 1873
  134. Genus Neopachycysta Hacker, 1928 accepted as Calotingis Drake, 1918
  135. Genus Octacysta Drake & Ruhoff, 1960 accepted as Dictyla Stål, 1874
  136. Genus Orthostira Fieber, 1861 (Misspelling)
  137. Genus Paraserenthia Wagner, 1940 accepted as Agramma Stephens, 1829
  138. Genus Phyllochisme Kirkaldy, 1904 accepted as Physatocheila Fieber, 1844
  139. Genus Phyllontochila Stål, 1873 (Misspelling)
  140. Genus Physacheila Kolenati, 1857 accepted as Physatocheila Fieber, 1844
  141. Genus Physamacheila Kolenati, 1857 accepted as Physatocheila Fieber, 1844
  142. Genus Physatochila Fieber, 1860 accepted as Physatocheila Fieber, 1844
  143. Genus Physodera Marshall, 1868 accepted as Derephysia Spinola, 1837 (unaccepted > junior objective synonym, unnecessary replacement name)
  144. Genus Prionostirina Schumacher, 1913 accepted as Urentius Distant, 1903
  145. Genus Serenthia Spinola, 1837 accepted as Agramma Stephens, 1829
  146. Genus Serenthiella Wagner, 1940 accepted as Agramma Stephens, 1829
  147. Genus Taphrostethus Fieber, 1844 accepted as Cantacader Amyot & Serville, 1843
  148. Genus Teratochila Drake & Poor, 1936 accepted as Pontanus Distant, 1902
  149. Genus Tropidochila Fieber, 1861 (Misspelling)
  150. Genus Wombalia Schouteden, 1919 accepted as Agramma Stephens, 1829
  151. Genus Abdastartus Distant, 1910 (uncertain, unassessed)
  152. Genus Aeipeplus Drake & Ruhoff, 1962 (uncertain, unassessed)
  153. Genus Aeithauma Drake & Ruhoff, 1965 (uncertain, unassessed)
  154. Genus Aeopelys Drake & Ruhoff, 1965 (uncertain, unassessed)
  155. Genus Aepykorys Drake & Ruhoff, 1965 (uncertain, unassessed)
  156. Genus Aframixia Drake & Ruhoff, 1960 (uncertain, unassessed)
  157. Genus Afrochila Rodrigues, 1980 (uncertain, unassessed)
  158. Genus Afrotingis Drake & Hill, 1964 (uncertain, unassessed)
  159. Genus Agachila Drake & Gómez-Menor, 1954 (uncertain, unassessed)
  160. Genus Agaotingis Drake, 1954 (uncertain, unassessed)
  161. Genus Aglotingis Drake, 1954 (uncertain, unassessed)
  162. Genus Aidoneus Distant, 1909 (uncertain, unassessed)
  163. Genus Alcletha Kirkaldy, 1900 (uncertain, unassessed)
  164. Genus Alinotingis Duarte Rodrigues, 1992 (uncertain, unassessed)
  165. Genus Alloecysta Drake, 1961 (uncertain, unassessed)
  166. Genus Ammianus Distant, 1903 (uncertain, unassessed)
  167. Genus Angiocader Drake, 1950 (uncertain, unassessed)
  168. Genus Angolotingis Drake, 1955 (uncertain, unassessed)
  169. Genus Angolusa Drake, 1958 (uncertain, unassessed)
  170. Genus Aphelotingis Drake, 1948 (uncertain, unassessed)
  171. Genus Archepopovia Golub, 2001 † (uncertain, unassessed)
  172. Genus Arushia Drake, 1951 (uncertain, unassessed)
  173. Genus Asperotingis Pericart, 2000 (uncertain, unassessed)
  174. Genus Astolphos Distant, 1904 (uncertain, unassessed)
  175. Genus Aulotingis Drake & Poor, 1943 (uncertain, unassessed)
  176. Genus Australocader Lis, 1997 (uncertain, unassessed)
  177. Genus Axiokersos Distant, 1909 (uncertain, unassessed)
  178. Genus Baichila Drake & Slater, 1956 (uncertain, unassessed)
  179. Genus Banahaona Drake & Ruhoff, 1961 (uncertain, unassessed)
  180. Genus Belenus Distant, 1909 (uncertain, unassessed)
  181. Genus Cadmilos Distant, 1909 (uncertain, unassessed)
  182. Genus Caffrocysta Duarte Rodrigues, 1981 (uncertain, unassessed)
  183. Genus Campylosteira Fieber, 1844 (uncertain, unassessed)
  184. Genus Cantacander (uncertain, unassessed)
  185. Genus Carvalhotingis Froeschner, 1995 (uncertain, unassessed)
  186. Genus Catoplatus Spinola, 1837 (uncertain, unassessed)
  187. Genus Cephalidiosus Guilbert, 1999 (uncertain, unassessed)
  188. Genus Cetiothucha Drake & Ruhoff, 1965 (uncertain, unassessed)
  189. Genus Chorotingioites Wappler, 2003 † (uncertain, unassessed)
  190. Genus Conchotingis Drake, 1954 (uncertain, unassessed)
  191. Genus Congochila Drake, 1954 (uncertain, unassessed)
  192. Genus Copium Thunberg, 1822 (uncertain, unassessed)
  193. Genus Corythauma Drake & Poor, 1939 (uncertain, unassessed)
  194. Genus Cysteocheila (uncertain, unassessed)
  195. Genus Daillea Pericart, 1991 (uncertain, unassessed)
  196. Genus Derephysia Spinola, 1837 (uncertain, unassessed)
  197. Genus Distocader Froeschner, 1968 (uncertain, unassessed)
  198. Genus Dolichocysta Champion, 1898 (uncertain, unassessed)
  199. Genus Engyotingis Drake & Ruhoff, 1961 (uncertain, unassessed)
  200. Genus Etesinalda Froeschner, 1996 (uncertain, unassessed)
  201. Genus Euahanes Distant, 1911 (uncertain, unassessed)
  202. Genus Exmesselensis Wappler, 2003 † (uncertain, unassessed)
  203. Genus Exulmus Froeschner, 1996 (uncertain, unassessed)
  204. Genus Froeschnerocader Pericart, 1986 (uncertain, unassessed)
  205. Genus Galeotingis Drake, 1947 (uncertain, unassessed)
  206. Genus Galotingis Duarte Rodrigues, 1980 (uncertain, unassessed)
  207. Genus Gitava Drake, 1948 (uncertain, unassessed)
  208. Genus Gonycentrum Bergroth, 1898 (uncertain, unassessed)
  209. Genus Gyalotingis Drake, 1960 (uncertain, unassessed)
  210. Genus Haedacanthus Duarte Rodrigues, 1992 (uncertain, unassessed)
  211. Genus Haedus Distant, 1904 (uncertain, unassessed)
  212. Genus Hebetingis Drake, 1960 (uncertain, unassessed)
  213. Genus Heissiella Pericart, 1984 (uncertain, unassessed)
  214. Genus Henrikus Drake, 1955 (uncertain, unassessed)
  215. Genus Holophygdon Kirkaldy, 1908 (uncertain, unassessed)
  216. Genus Hurdchila Drake, 1953 (uncertain, unassessed)
  217. Genus Hypsotingis Drake, 1960 (uncertain, unassessed)
  218. Genus Idiocysta China, 1930 (uncertain, unassessed)
  219. Genus Idiostyla Drake, 1946 (uncertain, unassessed)
  220. Genus Indocader Pericart, 1981 (uncertain, unassessed)
  221. Genus Intercader Golub & Popov, 1998 † (uncertain, unassessed)
  222. Genus Jingicoris Li & Zheng, 2002 (uncertain, unassessed)
  223. Genus Kalama Puton, 1876 (uncertain, unassessed)
  224. Genus Kapiriella Schouteden, 1919 (uncertain, unassessed)
  225. Genus Kibongoto Duarte Rodrigues, 1982 (uncertain, unassessed)
  226. Genus Larotingis Drake, 1960 (uncertain, unassessed)
  227. Genus Latitingis Pericart, 1985 (uncertain, unassessed)
  228. Genus Lemurichila Duarte Rodrigues, 1992 (uncertain, unassessed)
  229. Genus Leptobrysa (uncertain, unassessed)
  230. Genus Leptoptyx Drake & Ruhoff, 1965 (uncertain, unassessed)
  231. Genus Lullius Distant, 1904 (uncertain, unassessed)
  232. Genus Machairotingis Duarte Rodrigues, 1981 (uncertain, unassessed)
  233. Genus Macropharsa (uncertain, unassessed)
  234. Genus Madangocoris Pericart, 2000 (uncertain, unassessed)
  235. Genus Magmara Pericart, 1977 (uncertain, unassessed)
  236. Genus Malala Distant, 1910 (uncertain, unassessed)
  237. Genus Mcatella (uncertain, unassessed)
  238. Genus Metasalis Lee, 1971 (uncertain, unassessed)
  239. Genus Microcader Pericart, 1981 (uncertain, unassessed)
  240. Genus Miotingis Nel, 1992 † (uncertain, unassessed)
  241. Genus Monanthia Lepeletier & Audinet-Serville, 1828 (uncertain, unassessed)
  242. Genus Monosteira Costa, 1860 (uncertain, unassessed)
  243. Genus Naitingis Drake & Ruhoff, 1962 (uncertain, unassessed)
  244. Genus Naochila Drake, 1957 (uncertain, unassessed)
  245. Genus Neoangolusa Rodrigues, 1983 (uncertain, unassessed)
  246. Genus Neolasiotropis Wagner, 1961 (uncertain, unassessed)
  247. Genus Neoplerochila Duarte Rodrigues, 1981 (uncertain, unassessed)
  248. Genus Nesocypselas Kirkaldy, 1908 (uncertain, unassessed)
  249. Genus Nesocysta Kirkaldy, 1908 (uncertain, unassessed)
  250. Genus Nesotingis Drake, 1957 (uncertain, unassessed)
  251. Genus Oblongomorpha Wappler, 2003 † (uncertain, unassessed)
  252. Genus Oeocharis Drake & Ruhoff, 1965 (uncertain, unassessed)
  253. Genus Ogygotingis Drake, 1948 (uncertain, unassessed)
  254. Genus Oncochila Stål, 1873 (uncertain, unassessed)
  255. Genus Onymochita Drake, 1948 (uncertain, unassessed)
  256. Genus Oranoma Drake, 1951 (uncertain, unassessed)
  257. Genus Orotingis Drake & Poor, 1941 (uncertain, unassessed)
  258. Genus Orthosteira Fieber, 1844 (uncertain, unassessed)
  259. Genus Ottoicus Drake, 1960 (uncertain, unassessed)
  260. Genus Palauella Drake, 1956 (uncertain, unassessed)
  261. Genus Paleocader Froeschner, 1996 † (uncertain, unassessed)
  262. Genus Paraderephysia Pericart, 1978 (uncertain, unassessed)
  263. Genus Paragramma Lis, 2003 (uncertain, unassessed)
  264. Genus Paraleptoypha Duarte Rodrigues, 1980 (uncertain, unassessed)
  265. Genus Paraperissonemia Duarte Rodrigues, 1980 (uncertain, unassessed)
  266. Genus Paraphatnomella Lis, 2000 (uncertain, unassessed)
  267. Genus Parazetekella Nel, Waller & De Ploëg, 2004 † (uncertain, unassessed)
  268. Genus Perbrinckea Drake, 1956 (uncertain, unassessed)
  269. Genus Phatnocader Stusak, 1976 (uncertain, unassessed)
  270. Genus Phatnomella Pericart, 1981 (uncertain, unassessed)
  271. Genus Phyllontocheila Fieber, 1844 (uncertain, unassessed)
  272. Genus Placotingis Drake, 1960 (uncertain, unassessed)
  273. Genus Plerochila Drake, 1954 (uncertain, unassessed)
  274. Genus Plesionoma Drake, 1950 (uncertain, unassessed)
  275. Genus Pliobyrsa Drake & Hambleton, 1946 (uncertain, unassessed)
  276. Genus Pseudacalypta Pericart, 1983 (uncertain, unassessed)
  277. Genus Pseudohegesidemus Lis, 2001 (uncertain, unassessed)
  278. Genus Pseudurentius Pericart, 1992 (uncertain, unassessed)
  279. Genus Pullocader Pericart, 1991 (uncertain, unassessed)
  280. Genus Renaudea Drake, 1958 (uncertain, unassessed)
  281. Genus Sabestena Drake, 1944 (uncertain, unassessed)
  282. Genus Sanazarius Distant, 1904 (uncertain, unassessed)
  283. Genus Scymnotingis Drake, 1960 (uncertain, unassessed)
  284. Genus Sinalda Distant, 1904 (uncertain, unassessed)
  285. Genus Sisperonemia Rodrigues, 1987 (uncertain, unassessed)
  286. Genus Sphaerista Kirichenko, 1951 (uncertain, unassessed)
  287. Genus Stenotrachelus Guilbert, 1998 (uncertain, unassessed)
  288. Genus Stragulotingis Froeschner, 1969 (uncertain, unassessed)
  289. Genus Taichila Duarte Rodrigues, 1983 (uncertain, unassessed)
  290. Genus Taphnoma Pericart, 1991 (uncertain, unassessed)
  291. Genus Teleia Fieber, 1844 (uncertain, unassessed)
  292. Genus Teratocader Drake, 1950 (uncertain, unassessed)
  293. Genus Teratocheila (uncertain, unassessed)
  294. Genus Thaicader Pericart, 1991 (uncertain, unassessed)
  295. Genus Thaumamannia Drake & Davis, 1960 (uncertain, unassessed)
  296. Genus Therontingis Duarte-Rodrigues, 2002 (uncertain, unassessed)
  297. Genus Tingicader Golub & Popov, 1998 † (uncertain, unassessed)
  298. Genus Tropicotingis Duarte Rodrigues, 1981 (uncertain, unassessed)
  299. Genus Tropidocheila Fieber, 1844 (uncertain, unassessed)
  300. Genus Turenius Duarte Rodrigues, 1981 (uncertain, unassessed)
  301. Genus Ulmus Distant, 1905 (uncertain, unassessed)
  302. Genus Vatiga Drake & Hambleton, 1946 (uncertain, unassessed)
  303. Genus Xenotingis Drake, 1923 (uncertain, unassessed)
  304. Genus Ypsotingis Drake, 1947 (uncertain, unassessed)
  305. Genus Zeiratingis Drake & Ruhoff, 1961 (uncertain, unassessed)
  306. Genus Zelotingis Drake & Hambleton, 1946 (uncertain, unassessed)
  307. Genus Zorotingis Drake & Ruhoff, 1961 (uncertain, unassessed)
marine, brackish
recent + fossil
Not documented
IRMNG (2021). Tingidae Laporte, 1833. Accessed at: https://irmng.org/aphia.php?p=taxdetails&id=103421 on 2025-11-11
Date
action
by
2006-09-19 22:00:00Z
created
Rees, Tony
2011-12-31 23:00:00Z
changed
Rees, Tony

basis of record Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. [details] 

extant flag source Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. [details] 

habitat flag source Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. [details] 
Unreviewed
Classification Eukaryota (Superkingdom) > Animalia (Kingdom) > Eumetazoa (Subkingdom) > Arthropoda (Phylum) > Hexapoda (Subphylum) > Insecta (Class) > Pterygota (Subclass) > Neoptera (Infraclass) > Hemipterodea (=Oligonephridia) (Superorder) > Hemiptera (Order) > Cimicomorpha (Group) > Tingoidea (Superfamily) > Tingidae (Family) [details]

Descriptive info Bugs, terrestrial (Parker). Benton, 1993 habitat flag(s): T; stratigraphic range: Cretaceous (Berriasian)-Holocene (?Recent) [details]
To WoRMS 
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